Acompany that bred beagles for medical research has agreed to pay a record $35 million as part of a criminal plea, admitting it neglected thousands of dogs at its breeding facility in rural Virginia. Prosecutors said Monday that the penalties amount to the largest ever levied in an animal welfare case.

The plea deal also bars Envigo RMS and its West Lafayette, Indiana-based parent company, Inotiv, from breeding or selling dogs in the future.

Read the full story in the Richmond Times-Dispatch.